Ditch the wide stance, the running shoes, and the gloves.
The reason you don’t have enough ankle mobility is because of the fact that you are not allowing your torso to change angles toward the bottom of the lift. This is causing your knee to travel farther over your knee than it needs to in order to compensate, which in turn causes you to run out of ankle mobility toward the bottom of your lift.
Your feet are so far out that they are putting a huge torque on your femur, which is transferring all the way up to your hip. My hips ache just watching it haha. One of the things that you are doing wrong as well is that your feet are pointed way too far in. I know some guys recommend that kind of set up so that you can push out on the side wall of your shoe, but to me that just doesn’t seem optimal. It looks like you are just uncomfortable having your body in that contorted position. This is also probably the reason your knees cave slightly when you come up out of the hole.
In fact it looks like you are trying to stay perfectly upright the whole time, which is not possible if you are going to break parallel due to joint angles at the hip, and the circle that is drawn biomechanically around your hip via your femur. At some point, your torso will become more acute to the floor, which is not a terrible thing, because this will allow for your hips to come back in and move between your legs and give you that last bit of depth.
Correct all of these things and do paused squats to learn what it feels like to hit the bottom position and not be resting on your joints. (I mean seriously man, that has to make your hips sore as hell.) Once you fix it all up, your butt will be resting on your calves, and you can just hang out there all day and be comfortable. AND, if you time it right, you can get a little bounce to help you get the bar moving in the other direction.
When you can one rep somewhere between 300 and 365, buy a good belt and use it only for the top set of the day. This will also help you sit down more and help keep you upright if you don’t like falling forward at all.
Other than that, you seem to have made good progress. Not many people ever squat 250 anywhere close to depth.
Maybe someone can correct me if I am wrong so that I could learn as well though?